The correct answer is A. T.H. Morgan.
T.H. Morgan was an American geneticist who proved the idea of W.S. Sutton that genes are located on chromosomes. He did this by studying the inheritance of eye color in fruit flies. He found that the genes for eye color are located on the X chromosome, and that they are passed down from parents to offspring in a predictable way. This work helped to establish the field of genetics and to show that genes are located on chromosomes.
De Duve was a Belgian cytologist and biochemist who won the Nobel Prize in Physiology or Medicine in 1974 for his discovery of lysosomes. Palade was a Romanian-American cell biologist who won the Nobel Prize in Physiology or Medicine in 1974 for his discovery of ribosomes. Waldeyer was a German anatomist who coined the term “chromosome” in 1888.
De Duve, Palade, and Waldeyer all made important contributions to the field of biology, but they did not prove the idea of W.S. Sutton that genes are located on chromosomes. This was done by T.H. Morgan.